CRL: Mt Graham Red Squirrel Research Program- Publications and On-Going Research

On-Going and Future Research

As detailed throughout these web pages, the dynamic nature of the Pinaleño ecosystem requires frequent monitoring. These monitoring efforts chronicle fluctuations in squirrel abundance, food availability, vegetation composition, and weather. In addition, the Red Squirrel Monitoring Program (RSMP) is conducting basic and applied research on various aspects of the Pinaleño ecosystem. Current and future research projects will examine the impacts of introduced Abert's squirrels (Sciurus aberti) in the mountains and assess the mammalian community associated with the nutrient rich middens characteristic of red squirrels in coniferous forests. Furthermore, we endeavor to monitor survival, reproductive performance, and space use of the endangered red squirrels at a fine scale by carefully marking and observing squirrels from a distance and via radio telemetry. These data promise to provide critical insight on the demography and habitat use of Mt. Graham red squirrels in the ever-changing, high elevation forests of the Pinaleños. Revisit our web page for periodic updates on the status of these projects and to download reprints of research articles as they become available. Visit the KCRL conservation projects page for a list of current and past graduate students and their project descriptions.

Unpublished annual reports of the Mt. Graham red squirrel research project can be found here

Books -
Thorington, R. W., J. L. Koprowski, M. A. Steele, and J. Whatton. 2012. Squirrels of the World, Johns Hopkins University Press.
Sanderson, H. R., and J. L. Koprowski. 2009. The Last Refuge of the Mt. Graham Red Squirrel. University of Arizona Press, Tucson, AZ.
Steele, M. A., and J. L. Koprowski. 2001. North American Tree Squirrels. Smithsonian Institution Press: Washington, DC. 224 p.
Book Chapters -
Buecher, D.C., R. Sidner, J.L. Koprowski. 2010. Finding that 4-star diner or how bats might ‘anticipate’ productive foraging areas. Pp. 253-262 in: Halvorson, W., Schwalbe, C., Van Riper, C., III (eds.). Southwestern Desert Resources, University of Arizona Press.
Young, P. J. 1999. Red Squirrel, Tamiasciurus hudsonicus. In: Wilson, D.E., and S. Ruff, editors. The Smithsonian Book of North American Mammals. Smithsonian Institution Press: Washington, London. Pp. 460-461.
Young, P. J. 1994. Monitoring the Mount Graham Red Squirrel. In: Istock, C. A., and R. S. Hoffman,editors. Storm Over a Mountain Island. University of Arizona Press, Tucson. Pp. 227-246.
Book Reviews -
Chen, H.L., J.L. Koprowski. 2011. Book Review: Safe passages: highways, wildlife, and habitat connectivity. Journal of Mammalogy. 92: 1136-1141.
Peer-Reviewed Articles -
Abercrombie, S. T., J. L. Koprowski, M. H. Nichols, and J. S. Femhi. 2018. Native lagomorphs suppress grass establishment in a shrub-encroached, semiarid grassland. Ecology and Evolution. 9: 307-317. PDF of Article
Blount, S.J., J.L. Koprowski. 2012. Small mammal response to post-fire conditions: case of the endangered Mount Graham red squirrel. Southwestern Naturalist. 57:8-15. PDF of Article
Brown, D. E., R. D. Babb, C. Lorenzo, and M. M. Altemus. 2014. Ecology of the antelope jackrabbit (Lepus alleni). The Southwestern Naturalist 59:575-587. PDF of Article
Chen, H. L. and J. L. Koprowski. 2019. Can we use body size and road characteristics to anticipate barrier effects of roads in mammals? A meta-analyisis. Hystrix doi:10.4404/hystrix–00185-2018. PDF of Article
Chen, H. L. and J. L. Koprowski. 2016. Barrier effects of roads on an endangered forest obligate: influences of traffic, road edges, and gaps. Biological Conservation 199:33-40. PDF of Article
Chen, H. L. and J. L. Koprowski. 2015. Differential effects of roads and traffic on space use and movements of native forest-dependent and introduced edge-tolerant species. PLoS ONE 11:e0148121 doi:10.1371/journal.pone.0148121. PDF of Article
Chen, H. L. and J. L. Koprowski. 2015. Animal occurrence and space use change in the landscape of anthropogenic noise. Biological Conservation 192:315-322. PDF of Article
Chen, H. L. and J. L. Koprowski. 2013. Effects of roads on wildlife in Arizona: how far have we Traveled? In Gottfried, Gerald J.; Ffolliott, Peter F.; Gebow, Brooke S.; Eskew, Lane G.; Collins, Loa C., comps. 2013. Merging science and management in a rapidly changing world: Biodiversity and management of the Madrean Archipelago III and 7th Conference on Research and Resource Management in the Southwestern Deserts; 2012 May 1-5; Tucson, AZ. Proceedings. RMRS-P-67. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 593 p. PDF of Article
Cudworth, N. L., and J. L. Koprowski. 2014. Survival and mortality of the Arizona gray squirrel (Sciurus arizonensis). The Southwestern Naturalist 59:423-426. PDF of Article
Cudworth, N. L., and J. L. Koprowski. 2013. Foraging and reproductive behavior of Arizona gray squirrels (Sciurus arizonensis): impacts of climatic variation. Journal of Mammalogy 94:683-690. PDF of Article
Cudworth, N.L., J.L. Koprowski. 2011. Importance of scale in nest-site selection in Arizona gray squirrels. Journal of Wildlife Management. 75: 1668–1674. PDF of Article
Cudworth, N.L., J.L. Koprowski. 2010. Microtus californicus. Mammalian Species. 42: 230-243. PDF of Article
Cudworth, N. L., and J. L. Koprowski. 2010. First record of range extension of the cliff chipmunk (Tamias dorsalis) into the Huachuca Mountians. Western North American Naturalist 70:418-420. PDF of Article
Cudworth, N.L., J.L. Koprowski. 2010. Influences of mating strategy on space use of Arizona gray squirrels. Journal of Mammalogy. 91: 1235-1241. PDF of Article
Derbridge, J. J., E. E. Posthumus, H. L. Chen, and J. L. Koprowski. 2015. Solenodon paradoxus (Soricomorpha: Solenodontidae). Mammalian Species 47:100-106. PDF of Article
Derbridge, J. J., J. A. Merkle, M. E. Bucci, P. Callahan, J. L. Koprowski, J. L. Polfus, and P. R. Krausman. 2015. Experimentally derived δ13C and δ15N discrimination factors for gray wolves and the impact of prior information in Bayesian mixing models. PLoS ONE 10: e0119940. doi:10.1371/journal.pone.0119940. PDF of Aricle
Derbridge, J. J., P. R. Krausman, and C. T. Darimont. 2012. Using Bayesian stable isotope mixing models to estimate wolf diet ina multi-prey ecosystem. The Journal of Wildlife Managment 76:1277-1289. PDF of Article
Doumas, S. L., J. L. Koprowski, and W. O. Noble. 2016. Landscape-level assessment of Abert's squirrel and red squirrel in mixed conifer forest. The Southwestern Naturalist 60:240-246. PDF of Article
Doumas, S.L., J.L. Koprowski. 2013. Return of fire as a restoration tool: long-term effects of burn severity on habitat use by Mexican fox squirrels. Restoration Ecology 21:133-139. PDF of Article
Doumas, S. L., and J. L. Koprowski. 2013. Effect of heterogeneity in burn severity on Mexican fox squirrels following the retunr of fire. International Journal of Wildland Fire 22:405-413. PDF of Article
Edelman, A. J., J. L. Koprowski, and S. R. Bertelsen. 2009. Potential for nest site competition between native and exotic tree squirrels. Journal of Mammalogy 90: 167-174. PDF of Article
Edelman, A. J. and J. L. Koprowski. 2007. Communal nesting in asocial Abert's squirrels: the role of social thermoregulation and breeding strategy. Ethology 113: 147-154. Contact us for a PDF
Edelman, A. J. and J. L. Koprowski. 2006. Seasonal changes in home ranges of Abert's squirrels: Impact of mating season. Canadian Journal of Zoology 84: 404-411. PDF of Article
Edelman, A. J. and J. L. Koprowski. 2006. Characteristics of Abert's squirrel (Sciurus aberti) cavity nests. The Southwestern Naturalist 51(1): 64-70. PDF of Article
Edelman, A. J. and J. L. Koprowski. 2005. Selection of drey sites by Abert's squirrels in an introduced population. Journal of Mammalogy 86(6): 1220-1226. PDF of Article
Edelman, A. J., J. L. Koprowski, and J. L. Edelman. 2005. Kleptoparasitic behavior and species richness at Mt. Graham red squirrel middens. USDA Forest Service Proceedings RMRS-P-36: 395 - 398. PDF of Article
Edelman, A. J. and J. L. Koprowski. 2005. Diet and tree use of Abert's squirrels (Sciurus aberti) in a mixed-conifer forest. The Southwestern Naturalist 50(4): 461-465. PDF of Article
Erwin, J. A., K. Vargas, B. R. Blais, K. Bennett, J. Muldoon, S. Findysz, C. Christie, J. R. Heffelfinger, and M. Culver. 2019. Genetic assessment of a bighorn sheep population expansion in the Silver Bell Mountains, Arizona. Peer J 6:e5978. PDF of Article
Fitak, R. R., J. L. Koprowski, and M. Culver. 2013. Severe reduction in genetic variation in a montane isolate: the endangered Mount Graham red squirrel (Tamiasciurus hudsonicus grahamensis). Conservation Genetics 14:1233-1241. PDF of Article
Fulgham, K. M. and J. L. Koprowski. 2016. Kangaroo rat foraging in proximity to a colony of reintroduced black-tailed prairie dogs. The Southwestern Naturalist 61:194-202. PDF of Article
Goldstein, E. A., M. J. Merrick, and J. L. Koprowski. 2018. Low survival, high predation pressure present conservation challenges for an endangered endemic forest mammal. Biological Conservation 221:67-77. PDF of Article
Goldstein, E. A., M. J. Merrick, and J. L. Koprowski. 2017. Functional semelparity drives population dynamics and endangers a peripheral population. Biological Conservation 205:52-59. PDF of Article
González-Maya, J. F., R. Galindo-Tarazona, M. M. Urquijo Collazos, M. Zárate Vanegas, and A. Parra-Romero. Eds. 2017. El Oso Andino en el Macizo de Chingaza. Bogotá: Empresa de Acueducto, Alcantarillado y Aseo de Bogotá D. C.. EAB-ESP, Corporación Autónoma Regional del Guavio - CORPOGUAVIO, Parques Nacionales Naturales de Colombia (Parque nacional Natural Chingaza, Dirección Territorial Orinoquía) & Projecto de Conservación de Aguas y tierras - ProCAT Colombia. 120 Pp. PDF of Book
Gwinn, R. N. and J. L. Koprowksi. 2017. Differential response to fire by an introduced and an endemic species complicates endangered species conservation. Hystrix doi:10.4404/hystrix-27.2-11447:1-7. PDF of Article
Gwinn, R. N., J. L. Koprowski, Jessen, R., and M. Merrick. 2012. Sciurus spadiceus (Rodentia: Sciuridae). Mammalian Species 44:59-63. PDF of Article
Gwinn, R.N., G.H. Palmer, J.L. Koprowski. 2011. Sigmodon arizonae. Mammalian Species. 43: 149 – 154. PDF of Article
Gwinn, R.N., G.H. Palmer, J.L. Koprowski. 2011. Virginia opossum (Didelphis virginiana virginiana) from Yavapai County, Arizona. Western North American Naturalist. 71:113-114. PDF of Article
Hale, S. L., V. L. Greer,. J. L. Koprowski, and N. Ramos-Lara. 2018. Microsciurus santanderensis (Rodentia: Sciuridae). Mammalian Species 50:166-169. PDF of Article
Hale, S. L., and J. L. Koprowski. 2018. Ecosystem-level effects of keystone species reintroduction: a literature review. Restoration Ecology doi: 10.1111/rec.12684. PDF of Article
Hale, S. L., J. L. Koprowski, and H. Hicks. 2013. Review of black-tailed prairie dog reintroduction strategies and site selection: Arizona reintroduction. In Gottfried, Gerald J.; Ffolliott, Peter F.; Gebow, Brooke S.; Eskew, Lane G.; Collins, Loa C., comps. 2013. Merging science and management in a rapidly changing world: Biodiversity and management of the Madrean Archipelago III and 7th Conference on Research and Resource Management in the Southwestern Deserts; 2012 May 1-5; Tucson, AZ. Proceedings. RMRS-P-67. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 593 p. PDF of Article
Harrison, R. L., M. J. Patrick, and C. G. Schmitt. 2003. Foxes, fleas, and plague in New Mexico. The Southwestern Naturalist 48(4): 720- 722. PDF of Article
Hefty, K. L. and K. M. Stewart. 2018. Novel location data reveal spatiotemporal strategies used by a central-place forager. Journal of Mammalogy DOI:10.1093/jmammal/gyy019. PDF of Article
Hutton, K. A., J. L. Koprowski, V. L. Greer, M. I. Alanen, C. A. Schauffert and P. J. Young. 2002. Use of Spruce-Fir and Mixed-Conifer Forests by an Introduced Population of Abert's Squirrels (Sciurus Aberti). The Southwestern Naturalist 48: 257-260. PDF of Article

Jessen, T. G., A. K. Kilanowski, R. Nathan Gwinn, M. J. Merrick, and J. L. Koprowski. 2016. Microsciurus flaviventer (Rodentia: Sciuridae). Mammalian Species 48:59-65. PDF of Article

Jessen, R. R., G. H. Palmer, and J. L. Koprowski. 2013. Neotropical pygmy squirrels (Sciurillus pusillus) share termite nests. Ecotropica 19:73-76. PDF of Article
Jessen, R. R., R. N. Gwinn, and J. L. K0prowski. 2013. Sciurillus pusillus (Rodentia: Sciuridae). Mammalian Species 49:75-79. PDF of Article
Jessen, R. R., G. H. Palmer, and J. L. Koprowski. 2013. Maternity nest of an Amazon red squirrel in a bromeliad. Mastozoología Neotropical 20:159-161. PDF of Article
Jessen, R., M.J. Merrick, J.L. Koprowski, O. Ramirez. 2010. Presence of Guayaquil squirrels on the central coast of Peru: an apparent introduction. Mammalia. 74: 443-444. PDF of Article
Jo, Yeong-Seok, J. T. Baccus, and J. L. Koprowski. 2018. Mammals of Korea: a review of their taxonomy, distribution and conservation status. Zootaxa 4522: 001-216. PDF of Monograph
Jo, Yeong-Seok, J. J. Derbridge, J. T. Baccus. 2016. History and current status of invasive nutria and common muskrat in Korea. Wetlands DOI 10.1007/s13157-016-0867-z. PDF of Article
Kellam, J. O., D. K. Jansen, A. T. Johnson, R. W. Arwood, M. J. Merrick, and J. L. Koprowski. 2016. Big Cypress fox squirrel (Sciurus niger avicennia) ecology and habitat use in a cypress dome swamp-pine forest mosaic. Journal of Mammalogy 97:200-2010. PDF of Article
Ketcham, S. L., J. L. Koprowski, and D. A. Falk. 2017. Differential response of native Arizona gray squirrels and introduced Abert's squirrels to a mosaic of burn severities. Mammal Study 42:247-258. PDF of Article
Ketcham, S. L. and J. L. Koprowski. 2013. Impacts on wildfire on wildlife in Arizona: A synthesis. In Gottfried, Gerald J.; Ffolliott, Peter F.; Gebow, Brooke S.; Eskew, Lane G.; Collins, Loa C., comps. 2013. Merging science and management in a rapidly changing world: Biodiversity and management of the Madrean Archipelago III and 7th Conference on Research and Resource Management in the Southwestern Deserts; 2012 May 1-5; Tucson, AZ. Proceedings. RMRS-P-67. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 593 p. PDF of Article
Kilanowski, A. L. and J. L. Koprowski 2016. Female-biased sexual size dimorphism: ontogeny, seasonality, and fecundity of the cliff chipmunk (Tamias dorsalis). Journal of Mammalogy xx:x-x,DOI:10.1093/jmammal/gyw172. PDF of Article
Koprowski, J. L., M. A. Steele, and N. Ramos-Lara. 2016. Tamiasciurus mearnsi (Rodentia: Sciuridae). Mammalian Species 48:66-72. PDF of Article

Koprowski, J. L., L. Gavish, and S. L. Doumas. 2016. Sciurus anomalus (Rodentia: Sciuridae). Mammalian Species 48:48-58. PDF of Article

Koprowski, J. L., S. L. Doumas, M. J. Merrick, B. Oleson, E. E. Posthumus, T. G. Jessen, and R. N. Gwinn. 2013. It's lonely at the top: Biodiversity at risk to loss from climate change. In Gottfried, Gerald J.; Ffolliott, Peter F.; Gebow, Brooke S.; Eskew, Lane G.; Collins, Loa C., comps. 2013. Merging science and management in a rapidly changing world: Biodiversity and management of the Madrean Archipelago III and 7th Conference on Research and Resource Management in the Southwestern Deserts; 2012 May 1-5; Tucson, AZ. Proceedings. RMRS-P-67. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 593 p. PDF of Article
Koprowski, J. L. and R. Nandini. 2008. Global hotspots and knowledge gaps for tree and flying squirrels. Current Science 95: 851-856. PDF of Article
Koprowski, J. L., S. R. B. King, and M. J. Merrick. 2008. Expanded home ranges in a peripheral population: space use by endangered Mt. Graham red squirrels. Endangered Species Research, 4:227-232. Click here for online access
Koprowski, J. L., N. Ramos, B. S. Pasch, and C. A. Zugmeyer. 2006. Observations on the ecology of the endemic Mearns's squirrel (Tamiasciurus mearnsi). The Southwestern Naturalist 51(3): 426-430. PDF of Article
Koprowski, J. L., K. M. Leonard, C. A. Zugmeyer, and J. L. Jolley. 2006. Direct effects of fire on endangered Mount Graham red squirrels. The Southwestern Naturalist 51(1): 59-63. PDF of Article
Koprowski, J. L., M. I. Alanen, and A. M. Lynch. 2005. Nowhere to run and nowhere to hide: Response of endemic Mt. Graham red squirels to catastrophic forest damage. Biological Conservation 126: 491-498. PDF of Article
Koprowski, J. L. 2005. Annual cycles in body mass and reproduction of endangered Mt. Graham red squirrels. Journal of Mammalogy 86(2): 309-313. PDF of Article
Koprowski, J. L. 2005. Management and conservation of tree squirrels: The importance of endemism, species richness, and forest condition. USDA Forest Service Proceedings RMRS-P-36: 245-250.
PDF of Article
Koprowski, J. L., A. J. Edelman, B. S. Pasch, and D. C. Buecher. 2005. A dearth of data on the mammals of the Madrean archipelago: What we think we know and what we actually know. USDA Forest Service proceedings RMRS-P-36: 412- 415. PDF of Article
Koprowski, J. L. 2005. The response of tree squirrels to fragmentation: A review and synthesis. Animal Conservation 8: 369-376. PDF of Article
Koprowski, J. L. and M. C. Corse. 2005. Time budgets, activity periods, and behavior of mexican fox squirrels. Journal of Mammalogy 86(5): 947-952. PDF of Article

Koprowski, J. L. 2002. Handling tree squirrels with a safe and efficient restraint. Wildlife Society Bulletin 30: 101-103. PDF of Article (1.3 MB)
Koprowski, J. L. and M. C. Corse. 2001. Food habits of the Chiricahua fox squirrel (Sciurus nayaritensis chiricahuae). The Southwestern Naturalist 46(1): 62-65. PDF of Article
Koprowski, J. L. 1996. Natal philopatry, communal nesting, and kinship in fox squirrels and gray squirrels. Journal of Mammalogy 77(4): 1006-1016. PDF of Article
Koprowski, J. L. 1994. Sciurus niger. Mammalian Species 479: 1-9 . PDF of Article
Koprowski, J. L. 1994. Sciurus carolinensis. Mammalian Species 480: 1-9. PDF of Article
Koprowski, J. L. 1993. Alternative reproductive tactics in male eastern gray squirrels: "making the best of a bad job". Behavioral Ecology 4(2): 165-171. PDF of Article
Koprowski, J. L. 1993. The role of kinship in field interactions among juvenile gray squirrels (Sciurus carolinensis). Canadian Journal of Zoology 71: 224-226. PDF of Article
Koprowski, J. L. 1993. Behavioral tactics, dominance, and copulatory success among male fox squirrels. Ethology Ecology & Evolution 5: 169-176. PDF of Article
Koprowski, J. L. 1992. Removal of copulatory plugs by female tree squirrels. Journal of Mammalogy 73(3): 572-576. PDF of Article
Koprowski, J. L. 1991. Response of fox squirrels and gray squirrels to a late spring-early summer food shortage. Journal of Mammalogy 72(2): 367-372. PDF of Article
Koprowski, J. L. 1988. Longevity records for the fox squirrel. Journal of Mammalogy 69(2): 383-384 PDF of Article
Leonard, K. M. and J. L. Koprowski. 2009. A comparison of habitat use and demography of red squirrels at the southern edge of their range. American Midland Naturalist 162: 132-145. PDF of Article
Leonard, K. M. and J. L. Koprowski. 2009. Effects of fire on endangered Mt. Graham red squirrels (Tamiasciurus hudsonicus grahamensis): responses of individuals with known fates. Southwestern Naturalist 55:217-224. PDF of Article
Lurz, P. W. W., J. L. Koprowski, and D. J. Wood. 2008. The use of GIS and modelling approaches in squirrel population management and conservation: a review. Current Science 95(7): 918-922. PDF of Aricle
McColgin, M. E., J. L. Koprowski, and P. M Waser. 2017. White-nosed coatis in Arizona: tropical carnivores in a temperate environment. Journal of Mammalogy DOI: 10.1093/jmammal/gyx153. PDF of Article
McDonald, L. L., T. R. Stanley, D. L. Otis, D. E. Biggins, P. D. Stevens, J. L. Koprowski, W. Ballard. 2011. Recommended methods for range-wide monitoring of prairie dogs in the United States: U.S. Geological Survey Scientific Investigations Report. 2011–5063, 36 p.
Masoudi, Abolfazl, J. L. Koprowski, U. R. Bhattarai, and D. Wang. 2017. Elevational distribution and morphological attributes of the entomopathogenic fungi from forests of the Qinling Mountains in China. Applied Microbiology and Biotechnology. DOI: 10.1007/s00253-017-8651-4. PDF of Article
Merkle, J. A., J. L. Polfus, J. J. Derbridge, and K. S. Heinemeyer. 2017. Dietary niche partitioning among black bears, grizzly bears, and wolves in a multiprey ecosystem. Canadian Journal of Zoology 95663-671. PDF of Article
Merrick, M. J. and J. L. Koprowski 2017. Circuit theory to estimate natal dispersal routes and functional landscape connectivity for an endangered small mammal. Landscape Ecology DOI 10.1007/s10980-017-0521-z. PDF of Article
Merrick, M. J. and J. L. Koprowski 2017. Should we consider individual behavior differences in applied wildlife conservation studies? Biological Conservation 209: 34-44. PDF of Article
Merrick, M. J. and J. L. Koprowski 2016. Altered natal dispersal at the range periphery: the role of behavior, resources, and maternal condition. Ecology and Evolution 00:1-15. doi: 101002/ece3.2612. PDF of Article
Merrick M. J. and J. L. Koprowski 2016 Evidence of natal habitat preference induction within one habitat type. Proceedings of the Royal Society B 283: 20162106. PDF of Article
Merrick, M. J., S. L. Ketcham, and J. L. Koprowski. 2014. Sciurus ignitus (Rodentia: Sciuridae). Mammalian Species 46:93-100. PDF of Article
Merrick, M. J., J. L. Koprowski, and C. Wilcox. 2013. Into the third dimension: Benefits of incorporating LiDAR into wildlife habitat models. In Gottfried, Gerald J.; Ffolliott, Peter F.; Gebow, Brooke S.; Eskew, Lane G.; Collins, Loa C., comps. 2013. Merging science and management in a rapidly changing world: Biodiversity and management of the Madrean Archipelago III and 7th Conference on Research and Resource Management in the Southwestern Deserts; 2012 May 1-5; Tucson, AZ. Proceedings. RMRS-P-67. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 593 p. PDF of Article
Merrick, M. J., J. L. Koprowski, and R. N. Gwinn. 2012. Sciurus stramineus (Rodentia: Sciuridae). Mammalian Species 44: 44-50. PDF of Article
Merrick, M.J., J.L. Koprowski, R.N.Gwinn, G.H. Palmer, C.A. Zugmeyer. 2011. Surveys to determine the status of red squirrels in Guadalupe Mountains National Park, Texas. Southwestern Naturalist. 56: 24-28. PDF of Article
Merrick, M.J. R.N.Gwinn, R.L. Minor, R.R. Jessen, T.G. Jessen, V.L. Greer, J. L. Koprowski. 2010. Endangered Mount Graham Red Squirrel (Tamiasciurus hudsonicus grahamensis) uses nest following lightning strike. Southwestern Naturalist. 55: 123-124. PDF of Article
Merrick, M. J., S. R. Bertelsen, and J. L. Koprowski. 2007. Characteristics of Mount Graham red squirrel nest sites in a mixed conifer forest. The Journal of Wildlife Management 71: 1958-1963. PDF of Article
Minor, R. L. and J. L. Koprowski. 2015. Seed removal increased by scramble competition with an invasive species. PLoS ONE 10:e0143927. doi:10.1371/journal.pone.0143927 PDF of Article
Munroe, K. E., and J. L. Koprowski. 2014. Levels of social behaviors and genetic structure in a population of round-tailed ground squirrels (Xerospermophilus tereticaudus). Behavioral Ecology and Sociobiology 68:629-638. PDF of Article
Munroe, K.E., and J.L. Koprowski. 2012. Presence of copulatory plugs in round-tailed ground squirrels (Xerospermophilus tereticaudus). Southwestern Naturalist. 57: 208-210. PDF of Article
Munroe, K.E., and J.L. Koprowski. 2011. Sociality, Bateman’s gradients and the polygynandrous genetic mating system of round-tailed ground squirrels (Xerospermophilus tereticaudus). Behavioral Ecology and Sociobiology. 65: 1811-1824. PDF of Article
Nigenda-Morales, S. F., M. E. Gompper, D. Valenzuela-Galván, A. R. Lay, K. M. Kapheim, C. Hass, S. D. Booth-Binczik, G. A. Binczik, B. T. Hirsch, M. McColgin, J. L. Koprowski, K. McFadden, R. K. Wayne, and K. Koepfli. 2019. Phylogeographic and diversification patterns on the white-nosed coati (Nasua narica): Evidence for south-to-north colonization of North America. Molecular Phylogenetics and Evolution. 131: 149-163. PDF of Article
Palmer, G. H., J. L. Koprowski, and A. J. Pernas. 2014. Distribution and spread of an introduced insular population of red-bellied squirrels (Sciurus aureogaster) in Florida. Mammalia 78: 67-73. PDF of Article
Palmer, G. H., J. L. Koprowski, and A. J. Pernas. 2013. Nest tree and site selection of an introduced population of red-bellied squirrels (Sciurus aureogaster). Journal of Mammalogy 94:1274-1281. PDF of Article
Palmer, G. H., R. Nathan Gwinn, and J. E. Gwinn. 2009. Entanglement of a western screech-owl (Megascops kennicottii) in Coulter spiderling (Boerhaavia coulteri). The Southwestern Naturalist 54:523-524. PDF of Article
Palmer, G. H., J. L, Koprowski, and T. Pernas. 2008. Tree squirrels as invasive species: conservation and management implications. Pp 273 - 282 in G. L. Witmer, W. C. Pitt, K.A. Fagerstone (eds.), Managing vertebrate invasive species: proceedings of an international symposium. USDA/APHIS Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado. PDF of Article.
Palmer, R. R. and J. L. Koprowski. 2018. Influence of time and flood on diurnal mammal diversity and story level use in Igapó forest in the Peruvian Amazon. In: Myster R. (eds) Igapó (Black-water flooded forests) of the Amazon Basin. Springer, Cham. PDF of Chapter
Palmer, R. R. and J. L. Koprowski. 2017. Sciurus sanborni. Mammalian Species 49:93-96. PDF of Article
Palmer, R. R. and J. L. Koprowski. 2015. How do Neotropical pygmy squirrels (Sciurillus pusillus) use seasonally flooded forest in the Peruvian Amazon? Journal of Mammalogy DOI:10.1093/jmammal/gyv138. PDF of Article
Palmer, R. R., and J. L. Koprowski. 2014. Feeding behavior and activity patterns of Amazon red squirels. Mammalia 78: 303-313. PDF of Article
Parra-Romero, A., R. Galindo-Tarazona, J. F. González-Maya, and I. M. Vela-Vargas. 2019. Therya 10: DOI: 10.12933/therya-19-625 ISSN 2007-3364. PDF of Article
Pasch, B. and J. L. Koprowski. 2006. Sex differences in space use of Chiricahua fox squirrels. Journal of Mammalogy 87(2): 380-386. PDF of Article
Pasch, B. S. and J. L. Koprowski. 2005. Correlates of vulnerability in Chiricahua fox squirrels. USDA Forest Service proceedings RMRS-P-36: 426-428. PDF of Article
Pasch, B. S., J. L. Koprowski. 2011. Impacts of fire suppression on space use of Mexican fox squirrels. Journal of Mammalogy 92: 227-234 PDF of Article
Paudel, S., P. Pal, M. V. Cove, S. R. Jnawali, G. Abel, J. L. Koprowski, and R. Ranabhat. 2015. The endangered Ganges River dolphin Plantanista gangetica gangetica in Nepal: abundance, habitat and conservation threats. Endangered Species Research 29:59-68. PDF of Article
Posthumus, E. E., J. L. Koprowski, and R. J. Steidl. 2015. Red squirrel middens influence abundance but not diversity of other vertebrates. PLoS ONE 10(4):e0123633. doi:10.1371/journal/pone.0123633. PDF of Article
Ramos-Lara, N., and J. L. Koprowski. 2015. Spacing behavior of a non-larder-hoarding Tamiasciurus: a study of Mearns's squirrels in xeric coniferous forests. Ethology 121: 196-205. PDF of Article
Ramos-Lara, N., J. L. Koprowski, B. Kryštufek, and I. E. Hoffman. 2014. Spermophilus citellus (Rodentia: Sciuridae). Mammalian Species 46:71-87 PDF of Article
Ramos-Lara, N., J. L. Koprowski, and D. E. Swann. 2013. Nest-site characteristics of the montane endemic Mearns's squirrel (Tamiasciurus mearnsi): an obligate cavity-nester? Journal of Mammalogy 94:50-58. PDF of Article
Ramos-Lara, N., and J.L. Koprowski. 2012. Communal nesting behavior in Mearns’s squirrels (Tamiasciurus mearnsi). Southwestern Naturalist 57: 195-225. PDF of Article
Rushton, S. P., D. J. A. Wood, P. W. W. Lurz, and J. L. Koprowski. 2006. Modelling the population dynamics of the Mt. Graham red squirrel: Can we predict its future in a changing environment with multiple threats? Biological Conservation 131: 121-131. PDF of Article
Sanderson, H. R. and J. L. Koprowski, editors. 2009. The last refuge of the Mt. Graham red squirrel: ecology of endangerment. University of Arizona Press, Tucson. Order here!
Schauffert, C. A., J. L. Koprowski, V. L. Greer, M. I. Alanen, K. A. Hutton, and P. J. Young. 2002. Interactions Between Predators and Mt. Graham Red Squirrels (Tamiasciurus hudsonicus grahamensis). The Southwestern Naturalist 47: 498-501. PDF of Article
Scobie, E. H., R. R. Palmer, and J. L. Koprowski. 2014. Sciurus pyrrhinus (Rodentia: Sciuridae). Mammalian Species 46:88-92. PDF of Article
Steele, M. A., and J. L. Koprowski. 2001. North American Tree Squirrels. Smithsonian Institution Press: Washington, DC. 224 p.
Vela-Vargas, I. Mauricio, J. Sebastián Jiménez-Alvarado, Diego A. Zárrate-Charry, C. Moreno-Diaz, A. Parra-Romero, and J. F. González-Maya. 2017. Capítulo 3: Generalidades del Oso Andino (Tremarctos ornatus: Ursidae). In El Oso Andino en el Macizo de Chingaza. J. F. González-Maya, R. Galindo-Tarazona, M. M. Urquijo Collazos, M. Zárate Vanegas, and A. Parra-Romero Eds. Bogotá: Empresa de Acueducto, Alcantarillado y Aseo de Bogotá D. C.. EAB-ESP, Corporación Autónoma Regional del Guavio - CORPOGUAVIO, Parques Nacionales Naturales de Colombia (Parque nacional Natural Chingaza, Dirección Territorial Orinoquía) & Projecto de Conservación de Aguas y tierras - ProCAT Colombia. Pp. 58-120. PDF of Article
Wood, D. J. A., S. Drake, S. P. Rushton, D. Rautenkranz, P. W. Lurz, and J. L. Koprowski. 2007. Fine-scale analysis of Mount Graham red squirrel habitat following disturbance. The Journal of Wildlife Management 71(7): 2357-2364. PDF of Article
Wood, D. J. A., J. L. Koprowski, and P. W. W. Lurz. 2007. Tree squirrel introductions: A theoretical approach with population viability analysis. Journal of Mammalogy 88(5): 1271-1279. PDF of Article
Wood, D. J. A. 2007. Forest disturbance and the long term population persistence of the Mt. Graham red squirrel: a spatially explicit modeling approach. M. S. Thesis. University of Arizona, Tucson, AZ. 119pp. Contact John L. Koprowski for more information.

Young, P. J. 1999. Red Squirrel, Tamiasciurus hudsonicus. In: Wilson, D.E., and S. Ruff, editors. The Smithsonian Book of North American Mammals. Smithsonian Institution Press: Washington, London. Pp. 460-461.

Young, P. J. 1994. Monitoring the Mount Graham Red Squirrel. In: Istock, C. A., and R. S. Hoffman, editors. Storm Over a Mountain Island. University of Arizona Press, Tucson. Pp. 227-246.

Young, P. J., V. L. Greer, and S. K. Six. 2002. Characteristics of Bolus Nests of Red Squirrels in the Pinaleño and White Mountains of Arizona. The Southwestern Naturalist 47: 267-275. PDF of Article (1.5 MB)
Zugmeyer, C. A. and J. L. Koprowski. 2009. Severely insect-damaged forest: A temporary trap for red squirrels? Forest Ecology and Management 257: 464-470. PDF of Article
Zugmeyer, C. A. and J. L. Koprowski. 2009. Habitat selection is unaltered after severe insect infestation: Concerns for forest-dependent species. Journal of Mammalogy 90: 175-182. PDF of Article
Zugmeyer, C. A. and J. L. Koprowski. 2007. Avian nestling predation by endangered Mt. Graham red squirrel. The Southwestern Naturalist 52(1): 155-157. PDF of Article
Zugmeyer, C. A. 2007. Trailblazers in the forest: response of endangered Mt. Graham red squirrels to severe insect infestation. M.S. Thesis. University of Arizona, Tucson, AZ. 73 pp. PDF available through the U of A library, or contact John L. Koprowski for more information.

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