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CURRICULUM
VITAE
John
L. Koprowski
Wildlife
and Fisheries Science
School
of Natural Resources
214
Biological Sciences East
University
of Arizona
Tucson,
AZ 85721
Phone: (520) 626-5895 (Office) Fax: (520) 621-8801 E-mail: squirrel@ag.arizona.edu
Personal Data:
Born:
29 September 1961, Lakewood, OH
Married:
21 September 1985 with 2 children
Education:
University
of Kansas
Ph.D.(Honors) Biology, 1991, Postdoctoral Studies, 1991-92
Advisor: Kenneth B. Armitage
Dissertation:The
evolution of sociality in tree squirrels: the comparative behavioral ecology
of fox squirrels and
eastern gray squirrels.
Southern
Illinois University
M.A. Zoology, 1985
Advisor: W.D. Klimstra
Thesis:
Fox squirrel population trends and regulation in a southern Illinois woodlot,
1968-1984.
Ohio
State University
B.S. Zoology, 1983
Advisors: John Harder & Tony Peterle
Undergraduate
Research Thesis: Nocturnal activity of woodchucks (Marmota
monax) in an urban park
Honors and Awards:
2006
College of Agriculture and Life Sciences Diversity Award, University of
Arizona
Sabbatical, Fall 2007
Arizona Game and Fish Department Heritage Grant (2)
USFS Joint-Venture Agreement
Red Squirrel
Monitoring Program Grant. University of Arizona. Research Grant.
2005
President,
Arizona Chapter of The Wildlife Society
Outstanding Faculty Member, School of Natural Resources
National Park Service. Research Grant
Red Squirrel
Monitoring Program Grant. University of Arizona. Research Grant.
2004
President
Elect, Arizona Chapter of The Wildlife Society
College of Agriculture and Life Sciences Team Award, University of Arizona
Arizona Game and Fish Department. Research Grant
USFS
Joint-Venture Agreement. Research Grant
Desert
Southwest Cooperative Ecosystems Studies Unit. Research Grant.
Red Squirrel
Monitoring Program Grant. University of Arizona. Research Grant.
2003
Tenure
Granted
Outstanding
Faculty Member, School of Renewable Natural Resources
National
Park Service. Research Grant
Arizona
Game and Fish Department. 2 Research Grants.
USFS Joint-Venture
Agreement. Research Grant
Red Squirrel
Monitoring Program Grant. University of Arizona. Research Grant.
2002
Outstanding
Scholarly Achievement, School of Renewable Natural Resources
Arizona
Game and Fish Department Heritage Fund IIPAM. Research Grant
Red Squirrel
Monitoring Program Grant. University of Arizona. Research Grant.
Western
National Parks Association. Research Grant.
2001
USFS Joint-Venture
Agreement. Research Grant.
USFS Region
3 Species Assessment. Research Grant.
Desert
Southwest Cooperative Ecosystems Studies Unit. Research Grant.
Red Squirrel
Monitoring Program Grant. University of Arizona. Research Grant.
Scott Neotropical
Fund, Cleveland Zoological Society. Research Grant.
2000
Faculty Small Grant Program Award. University of Arizona. Research
Grant.
Red Squirrel Monitoring Program Grant. University of Arizona. Research
Grant.
Hewlett Award. Willamette University. Scientific Collections
Enhancement.
Faculty Achievement Award. Scholarship, Teaching, and Service.
Faculty Study Time Award, Willamette University. Competitive Course Release
1999
Emil W.
Haury Award for Outstanding Achievement in Science in the National Parks
Southwest
Parks and Monuments Association. Research Grant
Murdock
College Research Program for Life Sciences. Research Grant
Faculty
Achievement Award. Scholarship and Teaching, Willamette University
Atkinson
Award, Willamette University. Research Grant.
'Breakfast
with a Scientist', American Society of Mammalogists. Host
1998
Southwest
Parks and Monuments Association. Research Grant
Tenure
granted. Willamette University
National
Park Service Borderlands Program. Research Grant
1997
National
Geographic Society. Research Grant
National
Science Foundation-ILI (Co-PI). Instrumentation Grant
Southwest
Parks and Monuments Association. Research Grant
Promotion
to Associate Professor. Willamette University
1996
Southwest
Parks and Monuments Association. Research Grant
Murdock
Charitable Trust (Contributor). Institutional Research Grant
Faculty
Achievement Award for Scholarship, Teaching and Service. Willamette University
Junior
Sabbatical. Willamette University
1995
Southwest
Parks and Monuments Association. Two Research Grants
Faculty
21, Project Kaleidoscope
1994
National
Geographic Society. Research Grant
Southwest
Parks and Monuments Association. Research Grant
Atkinson
Award, Willamette University. Research Grant.
Courtesy
Assistant Professor of Fish and Wildlife. Oregon State University
Faculty
Achievement Award for Scholarship and Teaching, Willamette University
1993
Apprenticeships
in Science and Engineering. Research Grant (w/SR Kephart).
Atkinson
Award, Willamette University. Research Grant.
1991
The Dorothy
Haglund Outstanding Dissertation Award, The Graduate School, University
of
Kansas.
Dissertation
Honors, University of Kansas.
1990
The Graduate
Teaching Assistant of the Year, The Graduate School, University of Kansas.
University
of Kansas Graduate School Summer Fellowship
1988
University
of Kansas Graduate School Summer Fellowship
Sigma Xi,
The Scientific Research Society. Research Grant.
1986
Theodore
Roosevelt Memorial Fund, American Museum of Natural History. Research Grant.
Professional Societies:
American
Association for the Advancement of Science
American
Society of Mammalogists
Animal
Behavior Society
Council on Undergraduate Research
The
Ecological Society of America
International
Society for Behavioral Ecology
Sigma
Xi, The Scientific Research Society
Society
for Conservation Biology
Southwestern
Association of Naturalists
The
Wildlife Society
Publications (* designates undergraduate student author):
Books:
Steele, M. and J.L. Koprowski.
2001. North
American Tree Squirrels. Smithsonian Institution Press.
Sanderson, HR, Koprowski,
JL. (eds). Ecology of Endangerment: The Mt. Graham Red Squirrel and
its Last
Refuge (tentative title), University of Arizona Press.
Book Chapters and Journal Articles:
Koprowski, J.L., N. Rajamani.
2007. Global hotspots, centers of diversity, and conservation of the tree
and flying
squirrels. Current Science 92: In press.
Koprowski, J.L. 2007. Reproductive
strategies and alternative reproductive tactics of tree squirrels. In:
Wolff,
J. Sherman, P (eds). Rodent Societies. University of Chicago Press.
In press.
Lurz, P.W.W., J.L. Koprowski,
D.J.Wood. 2007. The use of GIS and modelling approaches in squirrel
population
management and conservation: a review. Current Science 92: In press.
Edelman, A.J., J.L. Koprowski.
2007. Communal nesting in asocial Abert’s squirrels: the role of
social
thermoregulation and breeding strategy. Ethology 113: 147-154. Contact
me for a PDF
Merrick, M., S.R. Bertelsen,
J.L. Koprowski. 2007. Nest site selection by endangered Mt. Graham red
squirrels.
Journal of Wildlife Management, 71: In press.
Edleman, A.J., J.L. Koprowski.
2007. Influence of female-biased sexual size dimorphism on dominance
of female
Townsend´s chipmunks. Canadian Journal of Zoology, 85: 1859-1863.
PDF
Johnson, H., P.R. Krausman,
V. Bleich, J.L. Koprowski. 2007. Effects of antler breakage on mating
Behavior
in male Tule elk (Cervus elaphus nannodes). European Journal
of Wildlife Research,
53: 9-15.
PDF
Wood, D.J., J.L. Koprowski,
P. Lurz. 2007. How small is too small? PVA assessment of tree squirrel
reintroduction
and ability to recover from small population sizes. Journal of Mammalogy
87: In
press.
Wood, D.J., S. Drake, S.P.
Rushton, D. Rautenkranz, P.W.W. Lurz and J.L. Koprowski. 2007. Fine
scale analysis
of Mount Graham red squirrel habitat following disturbance. Journal of
Wildlife
Management
71:In press.
Zugmeyer, C.A., J.L. Koprowski.
2007. Avian nest predation by endangered Mt. Graham red squirrels.
Southwestern
Naturalist, 52: In press.
Koprowski, J.L., R.J.Steidl.
2007. The dilemma of the small population: potential consequences of the
isolation
of Mt. Graham red squirrels. In: Sanderson, HR, Koprowski, JL. (eds).
Ecology of
Endangerment:
The Mt. Graham Red Squirrel and its Last Refuge (tentative title), University
of
Arizona
Press. Accepted in final form
Alanen, M I, Koprowski, J L,
Grinder M I, Greer, V L., Coates, C A, Hutton, K A. 2007. Habitat
characteristics
of the territories of Mt. Graham red squirrels: do sex differences exist?
In:
Sanderson,
HR, Koprowski, JL. (eds). Ecology of Endangerment: The Mt. Graham
Red Squirrel
and its
Last Refuge (tentative title), University of Arizona Press. Accepted
in final form.
Edelman, AE, JL Koprowski.
2007. Introduced Abert's squirrels on Mt. Graham: Could they be
contributing
to the decline of the red squirrel. In: Sanderson, HR, Koprowski, JL. (eds).
Ecology
of Endangerment:
The Mt. Graham Red Squirrel and its Last Refuge (tentative title), University
of Arizona
Press. Accepted in final form.
Greer, VL, JL Koprowski. 2007.
Time budget of the Mt. Graham red squirrel. In: Sanderson, HR,
Koprowski,
JL. (eds). Ecology of Endangerment: The Mt. Graham Red Squirrel and
its Last
Refuge
(tentative title), University of Arizona Press. Accepted in final form.
King, SRB, JL Koprowski.
2007. Effect of human and non-human disturbance on Mt. Graham Red
Squirrels.
In: Sanderson, HR, Koprowski, JL. (eds). Ecology of Endangerment:
The Mt.
Graham
Red Squirrel and its Last Refuge (tentative title), University of Arizona
Press. Accepted
in final
form.
Koprowski, J.L., T. Snow. 2007.
A comparison of the census methods used to monitor Mt. Graham red
squirrels.
In: Sanderson, HR, Koprowski, JL. (eds). Ecology of Endangerment:
The Mt. Graham
Red
Squirrel and its Last Refuge (tentative title), University of Arizona Press.
Accepted in final
form.
Munroe, K.E, J.L. Koprowski,
V.L. Greer. 2007. The social and mating systems of red squirrels: do Mt.
Graham
red squirrels fit the pattern? In: Sanderson, H.R, Koprowski, J.L. (eds).
Ecology of
Endangerment:
The Mt. Graham Red Squirrel and its Last Refuge (tentative title), University
of
Arizona
Press. Accepted in final form.
Bertelsen, S.R., J.L. Koprowski.
2007. Site characteristics of traditional and newly established middens
of
the Mount Graham red squirrel. In: Sanderson, H.R., Koprowski, J.L. (eds).
Ecology of
Endangerment:
The Mt. Graham Red Squirrel and its Last Refuge (tentative title), University
of
Arizona
Press. Accepted in final form.
Pasch, B.S., J.L. Koprowski.
2006. Annual cycles in body mass and reproduction of Chiricahua fox
squirrels
(Sciurus nayaritensis chiricahuae). Southwestern Naturalist,
51: In press.
Edelman, A.J., J.L. Koprowski.
2006. Seasonal changes in home ranges of Abert’s squirrels: impact of
mating
season and a novel habitat. Canadian Journal of Zoology, 84: 404-411. PDF
Koprowski, J.L., N. Ramos,
B.S. Pasch, C.A. Zugmeyer. 2006. Observations on the ecology of the
endemic
Mearns’s Squirrel (Tamiasciurus mearnsi). Southwestern Naturalist,
51:426-430. PDF
Rushton, S, D.J. Wood, P. Lurz,
J.L. Koprowski. 2006. Population viability analysis of the Mt. Graham
red squirrel:
can we predict its future in a changing environment? Biological Conservation,
131:
121-131.
PDF
Koprowski, J.L., K.M Leonard,
C.J. Zugmeyer, J.L. Jolley. 2006. Direct effects of fire on endangered
Mt. Graham
red squirrels. Southwestern Naturalist, 51: 59-63. PDF
Edelman, A.J., J.L. Koprowski.
2006. Characteristics of Abert’s squirrel (Sciurus aberti) cavity
nests.
Southwestern
Naturalist, 51: 64-70. PDF
Pasch, B. S., J. L. Koprowski.
2006. Sex differences in space use of Chiricahua fox squirrels.
Journal
of Mammalogy, 87: 380-386. PDF
Koprowski, JL. 2005. Annual
cycles in body mass and reproduction in the endangered Mt. Graham
red squirrel,
a montane island endemic. Journal of Mammalogy, 86:30-313. PDF
Koprowski, J.L., M.I. Alanen,
A.M. Lynch. 2005. Nowhere to run and nowhere to hide: response of
endemic
Mt. Graham red squirrels to catastrophic forest damage. Biological Conservation,
127:
491-498.
PDF
Koprowski, JL. 2005. Response
of tree squirrels to habitat fragmentation: a review and synthesis.
Animal
Conservation
8: 369-376. PDF
Koprowski, JL, MC Corse. 2005.
Activity, time budgets, and behavior of Mexican fox squirrels. Journal
of Mammalogy
86: 947-956. PDF
Edelman, A.J., J.L. Koprowski.
2005. Diet and tree use of Abert's squirrels (Sciurus aberti) in
a mixed-
conifer
forest. Southwestern Naturalist, 50: 461-465. PDF
Edelman, A.J., J.L. Koprowski.
2005. Selection of drey sites by Abert's squirrels in an introduced
population:
the role of structure versus tree species. Journal of Mammalogy 86:
1220-1226. PDF
Koprowski, J.L., G.T. Kellison,
S.L. Moneysmith. 2005. Status of red-bellied squirrels (Sciurus
aureogaster)
introduced to Elliott Key, Florida. Florida Field Naturalist, 33:
128-129.
Koprowski, JL. 2005. Management
and conservation of tree squirrels: the importance of endemism,
species
richness, and forest condition. Pp. 245-250 In Biodiversity and Management
of the
Madrean
Archipelago II. Proceedings RMRS-P-36. (Gottfried, G. J., Gebow, B. S.,
Eskew, L.
G., and
C. Edminster, eds.) United States Department of Agriculture, Forest Service,
Rocky
Mountain
Research Station, Fort Collins. PDF
Koprowski, J.L, A.J. Edelman,
B.S. Pasch, D.J. Buecher. 2005. A dearth of data on mammals in the
Madrean
Archipelago: what we think we know and what we do know. Pp. 413-415 In
Biodiversity
and Management of the Madrean Archipelago II. Proceedings RMRS-P-36.
(Gottfried,
G. J., Gebow, B. S., Eskew, L. G., and C. Edminster, eds.) United States
Department
of Agriculture,
Forest Service, Rocky Mountain Research Station, Fort Collins. PDF
Edelman, A.J., J.L. Koprowski,
J.L. Edelman. 2005. Kleptoparasitic behavior and species richness
at
Mt. Graham
red squirrel middens. Pp. 395-398. In Biodiversity and Management of the
Madrean
Archipelago
II. Proceedings RMRS-P-36. Gottfried, G. J., Gebow, B. S., Eskew, L. G.,
& Edminster,
C. (Eds).
RMRS-P-36. Fort Collins: United States Department of Agriculture, Forest
Service,
Rocky Mountain
Research Station. PDF
Pasch, B.S., J.L. Koprowski.
2005. Correlates of vulnerability in Chiricahua Fox Squirrels. Pp. 426-428.
In Biodiversity
and Management of the Madrean Archipelago II. Proceedings RMRS-P-36.
(Gottfried,
G. J., Gebow, B. S., Eskew, L. G., and C. Edminster, eds.) United States
Department
of Agriculture,
Forest Service, Rocky Mountain Research Station, Fort Collins. PDF
Koprowski, J.L. 2005 (July
20). Pine Squirrel (Tamiasciurus hudsonicus): a technical conservation
assessment.
[Online]. USDA Forest Service, Rocky Mountain Region. Available:
http://www.fs.fed.us/r2/projects/scp/assessments/pinesquirrel.pdf
McColgin, M., E. Brown, S.
Bickford, A. Eilers, J.Koprowski. 2003. Coati use of Century Plants.
Southwestern
Naturalist, 48:722-725. PDF
Hutton, K., J. Koprowski, V.
Greer, M. Alanen, Schauffert, C., P. Young. 2003. Use of spruce-fir
forests
by introduced Abert's squirrels. Southwestern Naturalist, 48:257-260. PDF
Schauffert, C., J. Koprowski,
V. Greer, M. Alanen, K. Hutton. 2002. Interactions between predators
and Mt.
Graham red squirrels. Southwestern Naturalist, 47:498-501. PDF
Koprowski, J.L. 2002. Handling
tree squirrels with an efficient and safe restraint. Wildlife
Society
Bulletin, 30:101-103. PDF
Koprowski, J.L. 2002. Book Review: An Atlas of Oregon Wildlife. Journal of Mammalogy 82.
Koprowski, J.L. 2002. Book Review: Animal Behavior. Journal of Mammalogy 82.
Koprowski, J.L., M.C. Corse.
2001. Food habits of Chiricahua Fox Squirrels (Sciurus
nayaritensis
chiricahuae). Southwestern Naturalist, 46:62-65. PDF
Koprowski, J.L., N. Perigo.
2000. Cooperative learning as a tool to teach vertebrate anatomy.
American
Biology Teacher, 26:282-284. PDF
Koprowski, J.L. 1998. Conflict
between the sexes: a review of social and mating systems of the
tree
squirrels. Pp. 33-41 in M.A. Steele, J.F. Merritt, D.A. Zegers (eds.),
Ecology and
Evolutionary
Biology of Tree Squirrels, Special Publication 6, Virginia Museum of
Natural
History. 310 pp.
Koprowski, J.L. and M.A. Steele.
1998. Future directions: Research on the ecology of tree
squirrels.
Pp. 309-310 in M.A. Steele, J.F. Merritt, D.A. Zegers (eds.), Ecology and
Evolutionary
Biology of Tree Squirrels, Special Publication 6, Virginia Museum of
Natural
History. 310 pp.
Koprowski, J.L. 1997.
Honing the craft of scientific writing: the role of peer review.
Journal
of College
Science Teaching, 27:133-135.
Koprowski, J.L. 1996. Natal
philopatry, communal nesting, and kinship in fox squirrels and
eastern
gray squirrels. Journal of Mammalogy, 77: 1006-1016. PDF
Koprowski, J.L. 1996. Book
Review of The Black-Tailed Prairie Dog. Journal of Wildlife
Management,
60:971-973.
Kneeland, M.C., J.L. Koprowski,
M.C. Corse. 1995. Potential predators of Chiricahua
fox
squirrels. Southwestern Naturalist, 40:340-342.
Koprowski, J.L. 1994. Sciurus carolinensis. Mammalian Species, 480:1-9. PDF
Koprowski, J.L. 1994. Sciurus niger. Mammalian Species, 479:1-9. PDF
Koprowski, J.L. 1993.
Alternative reproductive tactics in male eastern gray squirrels: "Making
the
best of a bad job". Behavioral Ecology 4:165-171. PDF
Koprowski, J.L. 1993.
Behavioral tactics, copulatory success, and dominance among male fox
squirrels.
Ethology Ecology & Evolution 5:169-176. PDF
Koprowski, J.L. 1993.
The role of kinship in field interactions of juvenile grey squirrels.
Canadian
Journal of Zoology 71:224-226. PDF
Koprowski, J.L. 1993.
Sex and species biases in scent marking by fox squirrels and eastern
gray
squirrels. Journal of Zoology (London) 230:319-323.
Koprowski, J.L. 1993.
Do estrous female gray squirrels, Sciurus carolinensis, advertise their
receptivity?
Canadian Field-Naturalist 106:392-394.
Fairbanks, L., J.L. Koprowski.
1992. Piscivory in fox squirrels. Prairie Naturalist
24:283-284.
Koprowski, J.L. 1992.
Removal of copulatory plugs by female tree squirrels. Journal of
Mammalogy
73:572-576. PDF
Koprowski, J.L. 1991.
Response of fox squirrels and eastern gray squirrels to a late
spring-early
summer food shortage. Journal of Mammalogy 72:367-372. PDF
Koprowski, J.L. 1991.
Damage due to scent marking by eastern gray and fox squirrels.
Proceeding
of the Great Plains Wildlife Damage Conference 10:101-105.
Koprowski, J.L. 1991.
Mixed-species mating chases of fox squirrels and eastern gray
squirrels.
Canadian Field- Naturalist 105:117-118.
Koprowski, J.L. 1991. Book Review: Food hoarding in animals. Journal of Insect Behavior 4.
Koprowski, J.L. 1989.
Burrow distribution of thirteen-lined ground squirrels in relation to tree
canopies.
Prairie Naturalist 21:185-188.
Koprowski, J.L. 1989.
Book Review: Reproductive success: studies of individual variation in
contrasting
breeding systems. Journal of Insect Behavior 2:449-450.
Koprowski, J.L., J.L. Roseberry,
W.D. Klimstra. 1988. Longevity records for the fox
squirrel.
Journal of Mammalogy 69:383-384. PDF
Koprowski, J.L. 1987.
Nocturnal activity of the woodchuck in an urban Ohio park. Canadian
Field-Naturalist
101:606-607.
Koprowski, J.L., N.M. Koprowski.
1987. Joint nest-building in the eastern gray squirrel.
Canadian
Field-Naturalist 101:610-611.
Koprowski, J.L. A. Nieto-Montes
de Oca, and R. M. Timm. Sciurus aureogaster. Mammalian
Species,
Accepted
pending minor revision.
Koprowski, J.L. A cost of larder
hoarding by acorn woodpeckers: theft by Chiricahua fox
squirrels.
Southwestern Naturalist, Accepted.
Koprowski, J. L., M. McColgin,*
and L. Compton. A handling cone for use on white-nosed coatis
(Carnivora:
Procyonidae) without chemical immobilants. Journal of Zoology, In
review.
Niedermeyer, J.*
and J. L. Koprowski. Home ranges of western gray squirrels (Sciurus
griseus) in a
mixed forest
fragment. Northwestern Naturalist, In review.
Robb, A.*
and J.L. Koprowski. Ecology of introduced eastern gray (Sciurus carolinensis)
and
native
western gray squirrels (S. griseus) in sympatry. Canadian Journal
of Zoology, In review.
Invited Seminars:
2006
Tree squirrels
as indicator species: forest condition and the conservation strategies.
4th International Conference on the Ecology of Tree Squirrels, India
(Plenary)
Conservation
of montane island endemics: the saga of the endangered Mt. Graham Red
Squirrel. Northern Arizona University
2005
Conservation
of montane island endemics: the saga of the endangered Mt. Graham Red
Squirrel. Iowa State University
Conservation
of montane island endemics: the saga of the endangered Mt. Graham Red
Squirrel. University of New Mexico (postponed)
Conservation
of tree squirrels: a North American perspective
9th International Mammalogical Congress, Sapporo, Japan
2003
Conservation of tree squirrels: future directions for the study of arboreal
squirrels
3rd International Conference on the Ecology of Tree Squirrels (Plenary)
The ecology of Mt. Graham red squirrels: trapped on a montane island.
New Mexico State University
2001
The ecology of Mt. Graham red squirrels: trapped on a montane island. Univ.
of Arizona
2000
The ecology of montane island mammals. Willamette University.
The ecology of montane island mammals. Univ. of Arizona
1997
Conflict
between the sexes: social and mating systems of tree squirrels. James
Cook Univ.
1996
Conflict
between the sexes: social and mating systems of tree squirrels. Bucknell
University.
Conflict
between the sexes: social and mating systems of tree squirrels. Baylor
University.
1995
Sociality
in tree squirrels: implications of sex differences and limited resources.
Univ. of
California, Davis.
1994
Social
ecology of tree squirrels: a comparative approach. Univ. Alberta.
Conflict
between the sexes: the evolution of social and mating systems in tree squirrels.
Concordia College.
Conflict
between the sexes: the evolution of social and mating systems in tree squirrels.
Univ.
Alaska Anchorage.
Intersexual
and intrasexual conflict: implications to sociality in tree squirrels.
Oregon State
Univ.
1993
Alternative
reproductive tactics of tree squirrels: intersexual and intrasexual conflict.
Univ. of
Oregon.
1992
Limited
resources: the desperate means of desperate squirrels. Willamette
University.
Conflict
between the sexes: social organization in tree squirrels. Trenton
State College.
Conflict
between the sexes: social organization in tree squirrels. Willamette
University.
Conflict
between the sexes: social organization in tree squirrels. Univ. of
Dallas.
1991
Conflict
between the sexes: social organization in tree squirrels. Emporia
State Univ.
1990
Behavioral
ecology of tree squirrels: individual & population-level consequences.
Louisiana
State Univ.
Papers Presented (since 1987):
2006
Global
hotspots, centers of diversity, and conservation of tree and flying squirrels.
(w/N. Rajamani). The Wildlife Society, Anchorage, AK.
Maternal
nesting behavior of the Mt. Graham red squirrel: Habitat and spatial influences.
(w/V.Greer). The Wildlife Society, Anchorage, AK.
Mortalities
in the endangered Mt. Graham red squirrel: Influence of predation and habitat
quality (w/M.Merrick). The Wildlife Society, Anchorage, AK.
Effects
of fire on endangered Mt. Graham red squirrel (w/S.Blount, D.Wood, T. Snow).
The
Wildlife Society, Anchorage, AK.
Resource
reduction related to insect infestation: Are Mt. Graham red squirrels
affected?
(w/C. Zugmeyer). American Society of Mammalogists, Amherst, MA.
Conservation
of tree and flying squirrels: world-wide challenges and solutions.
4th International Conference on the Ecology of Tree Squirrels, India
Home range
of endangered Mt. Graham red squirrels in insect damaged forests.
(w/C. Zugmeyer). 4th International Conference on the Ecology of Tree Squirrels,
India
The importance
of fire in the ecology of Mexican fox squirrels (w/B. Pasch). 4th International
Conference on the Ecology of Tree Squirrels, India
Functional
semelparity as a proximate cause of endangerment in the Mt. Graham red
squirrel
(w/V. Greer). 39th AZ-NM Chapter of The Wildlife Society, Flagstaff, AZ.
Nest site
selection of endangered Mt. Graham red squirrels in a mixed-conifer forest.
(w/M. Merrick, S. Bertelsen). 39th AZ-NM Chapter of The Wildlife Society,
Flagstaff, AZ.
Comparison
of nest-site characteristics between endangered Mt. Graham red squirrels
and
Introduced Abert’s squirrels (w/A.J. Edeman, S. Bertelsen). 39th AZ-NM
Chapter of The
Wildlife Society, Flagstaff, AZ.
Communal
nesting in the Mt. Graham red squirrel (w/V. Greer). 39th AZ-NM Chapter
of The
Wildlife Society, Flagstaff, AZ.
Predicting
midden locations based on habitat characteristics of the Mt. Graham red
squirrel
(Tamiasciurus hudsonicus grahamensis) (w/D.Wood, S.Rushton, P.Lurz, S.Drake).
39th
AZ-NM Chapter of The Wildlife Society, Flagstaff, AZ.
Resource
reduction related to insect infestation: Are Mt. Graham red squirrels
affected?
(w/C. Zugmeyer). 39th AZ-NM Chapter of The Wildlife Society, Flagstaff,
AZ.
Impact
of multiple insect outbreaks on Mt. Graham red squirrel habitat (w/A. Lynch,
M.
Alanen). 39th AZ-NM Chapter of The Wildlife Society, Flagstaff, AZ.
Ecology
of the Mearns’s squirrel (Tamiasciurus mearnsi) in the Sierra de San Pedro
Martir
(w/N. Ramos). 39th AZ-NM Chapter of The Wildlife Society, Flagstaff, AZ.
2005
Impacts
of forest fragmentation on tree squirrels: a comparative approach.
9th International
Mammalogical Congress, Sapporo, Japan.
Space use
and social organization of endangered Mt. Graham red squirrels: it's lonely
at the top.
(w/SRB King). The Wildlife Society, Madison, WI.
Direct
Effects of Fire on Endangered Mt. Graham Red Squirrels. (w/C. Zugmeyer,
K. Leonard,
J. Jolley). American Society of Mammalogists, Springfield, MO.
A PVA approach
to assess conservation strategies for the Mt. Graham Red Squirrel (w/D.
Wood). American Society of Mammalogists, Springfield, MO.
Comparative
Analysis of Home Range Size in the Endangered Mt. Graham Red Squirrel and
a
Closely Related Subspecies (w/K. Leonard). American Society of Mammalogists,
Springfield, MO.
Nesting
behavior of the Mount Graham red squirrel (Tamiasciurus hudsonicus
grahamensis) (w/V. Greer). American Society of Mammalogists, Springfield,
MO.
A
dearth of data on mammals in Arizona and New Mexico: what do we really
know? (w/A.
Edelman, B. Pasch, D. Buecher). AZ-NM Chapter of The Wildlife Society,
Gallup, NM.
Direct
Effects of Fire on Endangered Mt. Graham Red Squirrels. (w/C. Zugmeyer,
K. Leonard,
J. Jolley). AZ-NM Chapter of The Wildlife Society, Gallup, NM.
How
small is too small: Minimum viable population size in tree squirrels (w/D.
Wood)
AZ-NM Chapter of The Wildlife Society, Gallup, NM.
Comparative
Analysis of Home Range Size in the Endangered Mt. Graham Red Squirrel and
a
Closely Related Subspecies (w/K. Leonard) AZ-NM Chapter of The Wildlife
Society, Gallup, NM.
Cone
caching strategy and its impact on the Mt. Graham red squirrel population
(w/S. King,
S. Bertelsen). AZ-NM Chapter of The Wildlife Society, Gallup, NM.
Age
specific time budgets of the Mount Graham red squirrel (Tamiasciurus hudsonicus
grahamensis) (w/V. Greer). AZ-NM Chapter of The Wildlife Society, Gallup,
NM.
2004
Space use
and social organization of endangered Mt. Graham red squirrels: it's lonely
at the top.
American Society of Mammalogists, Arcata, CA (w/SRB King)
Influence
of sex on seasonal space use of Abert’s squirrels. American Society of
Mammalogists,
Arcata, CA (w/AJ Edelman)
Characteristics
of nest sites used by endangered Mount Graham red squirrels (Tamiasciurus
hudsonicus grahamensis). American Society of Mammalogists, Arcata, CA (w/SR
Bertelsen)
Cone caching
strategy and its impact on the Mt. Graham red squirrel population. American
Society of Mammalogists, Arcata, CA (w/S.R.B. King and SR Bertelsen)
Sex and
age differences in time budgets of endangered Mt. Graham red squirrels.
American
Society of Mammalogists, Arcata, CA (w/V Greer).
Space use
and social organization of endangered Mt. Graham red squirrels: it's lonely
at the
top. AZ-NM Chapter of The Wildlife Society, Safford AZ (w/SRB King)
Cone storage
in endangered Mt. Graham red squirrel middens: how the midden affects the
cone.
AZ-NM Chapter of The Wildlife Society, Safford AZ (w/M Yurenka)
Seeing
the forest for the trees: nest site selection of the endangered Mt. Graham
red squirrel.
AZ-NM Chapter of The Wildlife Society, Safford AZ (w/SR Bertelsen)
2004
Space use and social organization of endangered Mt. Graham red squirrels:
it's lonely at the top.
AZ-NM Chapter of The Wildlife Society, Safford AZ (w/SRB King)
Cone storage in endangered Mt. Graham red squirrel middens: how the midden
affects the cone.
AZ-NM Chapter of The Wildlife Society, Safford AZ (w/M Yurenka)
Seeing the forest for the trees: nest site selection of the endangered
Mt. Graham red squirre.
AZ-NM Chapter of The Wildlife Society, Safford AZ (w/SR Bertelsen)
2003
Impacts of fragmentation on tree squirrels: correlates, consequences, and
conservation. The Wildlife Society, Burlington.
Nesting ecology of introduced Abert's squirrels and endangered Mt. Graham
red squirrels (w/A.Edelman, S.Taylor). The Wildlife Society, Burlington.
Seeing the forest for the trees: nest-site selection of the endangered
Mount Graham red squirrels (Tamiasciurus hudsonicus grahamensis).
(w/S.Taylor). 3rd International Colloquium on Tree Squirrels. Newcastle-upon-Tyne,
England.
Space use of Chiricahua fox squirrels relative to fire history (w/BS Pasch)
How small is too small: minimum viable population size in tree squirrels
(w/D Wood)
Space use of sympatric endangered Mount Graham red squirrels and introduced
Abert’s squirrels (w/AJ Edelman)
Activity and Time Budgets of the Mt. Graham Red Squirrel (w/VL Greer)
A review
of methods for trapping, handling and marking tree squirrels with special
reference to
Mt. Graham red squirrels. Conference on the Ecology of the Mt. Graham
Red Squirrel, Safford, AZ.
A comparison
of the census methods used to monitor Mt. Graham red squirrels (w/T. Snow)
Conference on the Ecology of the Mt. Graham Red Squirrel, Safford, AZ.
The dilemma
of the small population: potential consequences of the isolation
of Mt. Graham
red squirrels (w/RJ Steidl). Conference on the Ecology of the Mt.
Graham Red Squirrel, Safford, AZ.
Habitat
characteristics of the territories of Mt. Graham red squirrels: do sex
differences exist? (w/Alanen,
M I, Grinder M I, Greer, V I, Coates, C A, and Hutton, K A). Conference
on the Ecology of the Mt. Graham
Red Squirrel, Safford, AZ.
Introduced
Abert's squirrels on Mt. Graham: Could they be contributing to the decline
of the red squirrel. (w/AJ Edelman)
Conference on the Ecology of the Mt. Graham Red Squirrel, Safford, AZ.
Effect
of human and non-human disturbance on Mt. Graham Red Squirrels. (w/SRB
King) Conference on the
Ecology of the Mt. Graham Red Squirrel, Safford, AZ.
The social
and mating systems of red squirrels: do Mt. Graham red squirrels fit the
pattern? (w/KE Munroe, VL Greer)
Conference on the Ecology of the Mt. Graham Red Squirrel, Safford, AZ.
Site characteristics
of traditional and newly established middens of the Mount Graham Red Squirrel.
(w/S Taylor)
Conference on the Ecology of the Mt. Graham Red Squirrel, Safford, AZ.
Impacts of fragmentation on tree squirrels: correlates, consequences, and
conservation.. American Society of Mammalogists, Lubbock.
Ecology of the endangered Mt. Graham Red Squirrel: a primer. AZ-NM Chapter
of The Wildlife Society, Gallup NM.
2002
Ecology of the social coati in semiarid montane islands: we're not in the
tropics anymore! Ecological Society of America, Tucson.
Response of endangered Mt. Graham red squirrels to insect infestations.
The Wildlife Society, Bismarck.
Ecology of a montane island endemic: Mexican fox squirrels. AZ-NM
Chapter of The Wildlife Society, Safford, AZ.
Response of endangered Mt. Graham red squirrels to insect infestations.
American Society of Mammalogists, Lake Charles, LA.
2001
Ecology of the social coati in semiarid montane islands: we're not in the
tropics anymore! The Wildlife Society, Reno.
Sex difference in the midden characteristics of Mt. Graham red squirrels
(w/M. Alanen, M. Grinder). The Wildlife Society, Reno.
Population trends of the Mount Graham red squirrel. (w/C. Schauffert,
V. Greer). The Wildlife Society, Reno.
Use of spruce-fir forest by introduced Abert's squirrels. (w/K. Hutton).
The Wildlife Society, Reno.
Sex differences in the ecology of Mexican fox squirrels. American
Society of Mammalogists, Missoula.
2000
Habitat
associations and microhabitat preferences in Chiricahua fox squirrels:
trapped on a sky
island (with M.Corse*, M. Kneeland*, N. Michel*). 2nd International Colloquium
on the
Ecology of Tree Squirrels.
Sex differences in time and activity budgets in Chiricahua fox squirrels
(with M. Corse*).
2nd International Colloquium on the Ecology of Tree Squirrels.
Sex and age differences in body fat composition in eastern gray squirrels
(with S. Howe*).
2nd International Colloquium on the Ecology of Tree Squirrels.
1999
Space use
by coatis: tropical carnivores in xeric habitat. American Society
of Mammalogists,
Seattle.
1998
Space use
by coatis in low-quality, semi-arid habitat. Conference of the Southwest
Parks and
Monuments Association, Tucson.
1996
Sociality
in tree squirrels: comparative ecology of tree and ground squirrels (Poster).
International Behavioral Ecology Conference, Canberra, Australia.
Ecology
of a montane island endemic: Chiricahua fox squirrels. Conference
of the Southwest
Parks and Monuments Association (Poster), Tucson.
1995
The ecology
of Chiricahua fox squirrels: trapped on a sky island (Poster).
Meeting of the
American Society of Mammalogists, Burlington, Vermont.
1994
Conflict
between the sexes: a review of social and mating systems of the tree squirrels.
International Colloquium on the Ecology of Tree Squirrels. Powdermill Biological
Station,
Pennsylvania
1993
Conflict
between the sexes: the mating systems of eastern gray squirrels and fox
squirrels
(Poster). Meeting of the American Society of Mammalogists, Bellingham,
Washington.
1991
Natal philopatry,
communal nesting, and kinship in tree squirrels: a comparative study.
Meeting of the American Society of Mammalogists, Manhattan, Kansas.
Kinship
in tree squirrel societies: don't leave home without it. Prairie
States Ecology
Conclave, Omaha, Nebraska.
1990
Social
organization of tree squirrels: potential management implications.
Midwest Fish &
Wildlife Conference, Minneapolis.
Why urban
wildlife research? The value of tree squirrels in an urban environment
(Poster)
National Symposium on Urban Wildlife, Cedar Rapids, IA.
Sex and
species-biased scent marking by fox squirrels and eastern gray squirrels.
Meeting of
the American Society of Mammalogists, Frostburg, MD.
1989
Scent marking
and bark-stripping damage by tree squirrels. Midwest Fish & Wildlife
Conference, Springfield, IL.
Alternative
reproductive tactics of male eastern gray squirrels. Meeting of the
American
Society of Mammalogists, Fairbanks.
1988
Critical
periods and the response of tree squirrels to natural variation in food
supply. Midwest
Fish and Wildlife Conference, Columbus, OH.
Alternative
male reproductive tactics in the eastern gray squirrel (Poster).
International
Behavioral Ecology Conference, Vancouver, Canada.
Mating
chases of the fox and gray squirrel. Prairie States Ecology Conclave,
Eureka Springs,
AR.
Professional Service:
Reviewer for Science,
Ecology, Biological Conservation, Animal Behaviour, Ethology, Oecologia,
Journal
of Mammalogy, Mammalian Species, American Midland Naturalist, The Southwestern
Naturalist,
Journal of Wildlife Management, Behavioral Ecology and Sociobiology, Behavioral
Ecology,
The Canadian Field-Naturalist, Journal of College Science Teaching, The
Prairie
Naturalist,
Acta Theriologica, Academic Press, WC Brown/McGraw-Hill, National Science
Foundation,
Mammalia, Wildlife Monographs, Journal of Applied Ecology, Western North
American
Naturalist, Zoo Biology, Journal of the Pennsylvania Academy of Sciences,
Revue d'écologie
: la terre et la vie, Tropical Zoology, Current Science
Member, Vertebrate Species
Team, Sonoran Desert Conservation Plan (2006 to present)
Member, Thick-billed Parrot
Reintroduction Team, Arizona Game & Fish Dept. (2006 to present)
Member, Technical Subteam,
Mt. Graham Red Squirrel Recovery Team (2002 to present)
Technical Advisor to Western
Gray Squirrel Recovery Team, Washington Dept. of Fish & Wildlife (2003)
Grants and Awards Committee,
American Society of Mammalogists (2002 to present)
Aldo Leopold Conservation
Award Committee, American Society of Mammalogists (2003 to present)
Organizing Committee, 3rd
International Colloquium on the Ecology of Tree Squirrels, Newcastle-upon-Tyne
(2003)
Host and Organizer, 2nd International
Colloquium on the Ecology of Tree Squirrels, Salem, OR (2000)
President, Arizona Chapter
of The Wildlife Society (2004-2006)
Board Member, Arizona Chapter
of The Wildlife Society (2006 to present)
Student Presentation Evaluator,
American Society of Mammalogists (1999-2003)
Vice-President, Society for
Northwestern Vertebrate Biology (1993)
Member of the Lagomorph/Sciurid
Strategic Planning Team, Kansas Dept.of Wildlife and Parks
(1988).
University Service:
Director, Desert Southwest
Cooperative Ecosystems Studies Unit (Nov 2002-2003)
SRNR Faculty Status Committee
(2003 to 2006, chair 2005-06)
RNR Admissions Committee (2002
to present)
SRNR Seminar Series Coordinator.
Spring 2001.
School of Renewable Natural
Resources Awards Committee (2001-present).
The Willamette University
Faculty Council (1999-2000). Elected by peers.
Undergraduate Research Committee
(1994-1996, 1998-1999; Chair 1995-96)
International Education Committee
(1997-1998)
University Representative
to the Malheur Field Station Board (Fall 1992 to present)
Library Advisory Committee
(Fall 1994 to present)
Search Committees: Psychology
and Exercise Science (1993), Serials Librarian (1994)
Undergraduate Awards Assistant (1996), Environmental Science (1996), Forestry
Extension (2002), Watershed Faculty (2 positions:2005), SNR Director
(2006)
Saeculum Vocat, Junior Career
Session (1998, 1999)
Biology Club Advisor (1993-94)
Presentations (4) to Willamette
Outdoors and Residence Halls
Presentations (3) to Board
of Trustees (1993, 1994 and 1998)
Community Service:
Elementary School Visits, Quail
Run Elementary (5 from 2001 to present)
Middle School Visits, Tortolita
Middle School (2 from 2004 to present)
Endangered Species Panelist,
Waldo Middle School (1996 to 2000)
Leader, YMCA Day Care tour
of Willamette (1995)
Campfire talks (8) for the
National Park Service (1994 to 1999)
Host, job shadowers from local
schools (9 from 1993-1999)
4-H Spring Field Day (1993)
Media Collaborations:
Guest, Birds & Nature,
1330 KPTT Pittsburgh, 45 minute segment
Host,
Creature of the Week Discussion, Discovery Channel On-line, 2000
Collaborator
on pitch tape with National Geographic TV, May-June 1999
Featured
on The World of the Chiricahua Mountains, New Mexico State University,
1998
Featured
on Oregon Field Guide, Oregon Public Broadcasting, 1996
Scientific
Adviser:
The
Gray Squirrel, Morningstar Productions, 1999
Tall
Tales from a Mountain Slope/Los Rabos Altos de la Ladera by Maria L.
Retana, High Desert Productions, Bisbee.
Ecology of the Endangered
Mt. Graham Red Squirrel (Tamiasciurus hudsonicus grahamensis)
Dates:
2000
to present
Location:
Pinaleno Mountains and White Mountains of Arizona
Project
Description: The Mt. Graham red squirrels is an endangered subspecies
of the widespread red squirrel. Found only in the Pinaleno
(AKA Graham) Mountains, the Mt. Graham red squirrel was once thought
to have been extirpated (as recently as the late 1960's) from these mountains.
However, subsequent reports verified the existence of a small population
in the high elevation forests of the Pinalenos. After listing as
an endangered species in the 1980's, the species has been monitored through
regularly quarterly censuses by the University of Arizona's Red Squirrel
Monitoring Program. Presently, we continue this monitoring effort
and are invoking on several studies to examine various aspects of red squirrel
ecology. Currently, we are conducting intensive live trapping and
radiotelemetric analyses of endangered Mt. Graham red squirrels to learn
more about their space use, reproductive tactics and success, demographics,
and habitat use. Meghan
Yurenka, an M.S. student, is examining the microclimate of occupied
and unoccupied middens to test the idea that middens are cool, moist places
to store cones. Kate Leonard, an M.S. student, is the investigating
the impacts of experimental fuel wood removal strategies on the demography
of Mt. Graham red squirrels to assess how fire management and squirrel
management strategies intersect. Claire
Zugmeyer, an M.S. student, is working on the ecology of MGRS in high
elevation spruce-fir forests that have been severely damaged by insects
to discover just how a few MGRS have been able to cope with the massive
habitat changes induced by insects. Finally, David Wood, an M.S.
student, is endeavoring to link the demography of MGRS collected by radiotelemetric
analysis with spatial data from high resolution imagery in a collaborative
venture with the UA's Office of Arid Lands and the Univ of Newcastle-upon-Tyne's
Centre for Life Science Modeling.
Ecology of Introduced
Abert's Squirrels (Sciurus aberti) on Mt.Graham
Dates:
2001
to present
Location:
Pinaleno Mountains of Arizona
Project
Description: Abert's squirrels were introduced to the Pinaleno Mountains
of Arizona in the early 1940's by the Arizona Department of Game and Fish
in an effort to enhance hunting opportunities for the populace, a common
conservation practice of the time. By the 1960's, biologists began
to note the potential negative impact of this introduction on the native
Mt. Graham red squirrel. Despite these concerns, there have been
no published studies on the potential ecological overlap between these
species. Abert's squirrels are often describes as Ponderosa pine
obligates; however, we have good evidence that on Mt. Graham the Abert's
squirrels are resident in areas that are depauperate of pine. Andrew
Edelman, an M.S. student, examined the space and habitat use of Abert's
squirrels using radiotelemetry on Mt. Graham and Nate
Gwinn, an M.S. student, is examining the response of Abert's squirrels
to fire induced habitat changes. Current hopes are to initiate an
Abert's squirrel removal experiment to assess the role of this exotic species
in the decline of endangered Mt. Graham red squirrels.
Ecology of the Arizona
Gray Squirrel: an unknown native species AND
Status of the Catalina
Gray Squirrel: The implications of introduced Abert's Squirrels
Dates:
2000
to
present
Location:
Huachuca Mountains, Catalina and Rincon Mountains of Arizona
Project
Description: Just as in the previous research project, Abert's squirrels
were introduced into the Catalina Mountains just outside of Tucson in the
early 1940's. Sometime after they appear to have colonized the adjacent
Rincon Mountains. In the 1960's concern was expressed over the status
of the Catalina Gray Squirrel, a subspecies of the Arizona Gray Squirrel.
Few records have been reported in the Catalina Mountains since this time.
Currently, we are revisiting sites where museum specimens have been collected
for each species and using hair collection tubes and observations, attempting
to document the continued coexistence of both species. Studies in the adjacent
Rincon Mountains suggest that Abert's squirrels now dominate the mountains
while the native Arizona Gray Squirrel may be relegated to isolated areas.
Nichole
Cudworth, MS student, has recently extended our efforts to the Huachuca
Mountains has initiated live trapping and radiotelemetric studies.
Ecology of Introduced Mexican Red-bellied Squirrels in Biscayne National Park and their impact on an Endangered Palm
Two pairs of Mexican Red-bellied Squirrels (Sciurus aureogaster) were introduced to the 10 km2 Elliott Key in Biscayne National Park, Florida, USA about 70 years ago. The population expanded to the point where it was believed to negatively impact an endangered endemic palm. In 1992, the 7 m tidal surge that accompanied Hurricane Andrew submersed the entire key and the introduced tree squirrel was believed to have been eradicated as a result. Recent anecdotal observations suggested that the population was extant and growing. A visit by one of us (John Koprowski) to the site in March 2005 documented the persistence of the population. Our study will investigate the distribution, habitat use, and impact on palms of Mexican red-bellied squirrels using drey surveys, live trapping, radiotelemetry, and observation. Field work will be initiated in December 2005 and continue for 2 years. Geoff Palmer, an MS student, continues to document the ecology of the squirrel and its potential impact on the Sargent's Palm.
The conservation and ecology of Mearns's Squirrel (Tamiasciurus mearnsi) in Sierra San Pedro Martir, Baja California, Mexico
The Mearns's squirrel is an
montane island endemic found in the highest regions of Baja California's
greatest mountain range at elevations around 3000 m. The species is considered
to be threatened in Mexico and no ecological studies have been conducted.
In addition, eastern gray squirrels (Sciurus carolinensis) were
introduced to lower elevations over 50 years ago and their status is unknown.
Related species in the United States typically larderhoard conifer cones.
In our first visit to the region in November 2004, we caught glimpses of
only two animals and did not find any larderhoards suggesting the behavior
and ecology of this species may be quite interesting. Field efforts began
spring 2005 the capture and radiocollaring of animals to assess their habitat
use and basic ecology. Nicolas
Ramos, a PhD student, will continue this fieldwork into 2008.
Conservation and
behavioral ecology of the rare Chiricahua fox squirrel
Dates:
1993
to present
Location:
Chiricahua
National Monument, Arizona.
Project
Description: A Master's student, Bret
Pasch, recently completed work on this project incorporating radiotelemetry
to examine space and habitat use of this uncommon species. Sandy
Doumas, an MS student, is expanding our research to examine the entire
Chiricahua Mountain range to assess the use of fire-impacted sites by Mexican
fox squirrels. The mountains of the desert southwest function as relatively
cool lush islands of forested habitat in a sea of desert. The Chiricahua
Mountains in extreme southeastern Arizona function as one such island and
are unique in their location at the southern terminus of the Rocky Mountains
and the northern extent of the Sierra Madre. As a result of this
unique location, the diversity of the Chiricahua montane (or sky) island
is great. Chiricahua fox squirrels (Sciurus nayaritensis
chiricahuae) are a large bodied subspecies of the Mexican fox squirrel
(S. nayaritensis) found only in the Chiricahuas. Ecological
data on Chiricahua fox squirrels is scant and resulted in a listing as
a Category 2 species by the US Fish and Wildlife Service (a category that
was eliminated by the USFWS). Our study represents the first ecological
study of this montane isolate. Our efforts focused primarily on habitat
use, nest tree selection, activity/movement patterns, and reproductive
performance. However, the large size of this tree squirrel allowed
us to examine current ideas on the evolution of sociality in squirrels
in relation to body size. More than 2 dozen undergraduate students
have worked on this research project with me over the years and several
have produced senior theses from ecological investigations:
Habitat use and nest site location of Chiricahua fox squirrels--Nicole
Michel
Foraging behavior of male and female Chiricahua fox squirrels--Michele
Corse
Vigilance behavior of Chiricahua fox squirrels in relation to human activity--Mary
Kneeland
A Comparison of
Habitat between Sites Occupied by Black-tailed Prairie Dogs in Sonora and
Unoccupied Sites in Arizona
Dates:
2001
to 2004
Location:
Grasslands of SE Arizona and Northern Sonora
Project
Description: Black-tailed prairie dogs were extirpated from Arizona
early in the 20th century. BT prairie dog populations appear to be
declining significantly throughout their range. Recently, several
relictual populations were rediscovered in northern Sonora just south of
Arizona. Carol Coates, an M.S. student, examined characteristics
of these relictual sites with those of sites in the high elevation grasslands
of southeastern Arizona.
Juvenile body condition
as a proximate mechanism of dispersal and social group formation in tree
squirrel
Dates:
1999
to 2001
Location:
Cascade and Coast Ranges of Oregon; Salem, Oregon
Project
Description: This project is a new venture scheduled for Summer
1999. Group formation in mammals often involves the retention of
young in their natal area. This study examines the hypothesis that
body condition is important in the natal dispersal of juvenile eastern
gray squirrels (Sciurus carolinensis), a highly social species,
and western gray squirrels (Sciurus griseus), a solitary species.
Western gray squirrels are a threatened species in the State of Washington
but are not of conservation concern in Oregon. The body condition
of young will be experimentally enhanced through the provision of supplemental
food. The impact of body condition on the rates of social interaction,
group cohesiveness, and dispersal from the birthsite will be examined.
Two undergraduates worked with me during the first year of this project,
Scott Howe and Matt Woolsey. During the summer of 2000, three undergraduates,
Ben Hudson, Jason Niedermeyer, and Aaron Tsuha conducted independent research
on the project.
Click HERE
for a research update on this project
Ecology and conservation
of a social carnivore: sociality of coatis in a xeric environment.
Dates: 1995 to present
Location:
Chiricahua
National Monument, Arizona
Project
Description: Social systems often vary
dependent on the distribution of resources. White-nosed coatis (Nasua
narica) are diurnal social carnivores related to raccoons that range
from northern South America to the southwestern deserts of the United States,
where they are a species of special concern. The social system of
this species is unique amongst the Carnivora for the larger bodied males
are solitary while the smaller females form groups of 10-20 animals.
Evidence from tropical forests suggests that males range over greater areas
than females. The arid mountains of the Arizona deserts provide a
vastly different environment with resources much less common and more widely
dispersed than in tropical forests. The objective of this project
is to examine the plasticity of social behavior and space use in coatis
by examining their ecology in aridlands. Thusfar, we have documented
that home ranges are often 10X greater than those in the tropics and that
males and females range equally widely (or during some seasons females
range MORE widely than males). This project has involved more than
a dozen undergraduate students, most of whom have completed independent
research projects or senior theses on ecological topics using coatis as
a model. Presently, I am collaborating with Maureen McColgin, a Ph.D.
student at Purdue University who is examining the mating system of these
coatis using molecular techniques. Past undergraduate student research
topics include:
Population
Ecology:
Differential habitat use of male and female coatis--Robin
Panzer
Influence of seed and lizard availability on space use patterns--Erica
Brown
Arthropod availability as a determinant of space use in male and female
coatis--Ian Silvernail
Impacts of El Nino on home range size in male and female coatis--Jessica
Girard
Behavioral
Ecology:
Differential space use and patterns of spatial overlap of male and female
coatis--Sophia Bickford
Den site selection of male and female coatis--Maureen
McColgin
Postparturition group formation in adult female coatis--Anna
Eilers
Conservation:
Impacts of fire on habitat use by coatis--Suzanne
Piluso
Influence
of human activity on coatis in a National Monument--Jed
Jorgenson
Conservation and
behavioral ecology of the rare Chiricahua fox squirrel
Dates:
1993
to present
Location:
Chiricahua
National Monument, Arizona.
Project
Description: A Master's student, Bret
Pasch, is working on this project incorporating radiotelemetry to examine
space and habitat use of this uncommon species. The mountains of the desert
southwest function as relatively cool lush islands of forested habitat
in a sea of desert. The Chiricahua Mountains in extreme southeastern
Arizona function as one such island and are unique in their location at
the southern terminus of the Rocky Mountains and the northern extent of
the Sierra Madre. As a result of this unique location, the diversity
of the Chiricahua montane (or sky) island is great. Chiricahua
fox squirrels (Sciurus nayaritensis chiricahuae) are a large bodied
subspecies of the Mexican fox squirrel (S. nayaritensis) found only
in the Chiricahuas. Ecological data on Chiricahua fox squirrels is
scant and resulted in a listing as a Category 2 species by the US Fish
and Wildlife Service (a category that was eliminated by the USFWS).
Our study represents the first ecological study of this montane isolate.
Our efforts focused primarily on habitat use, nest tree selection, activity/movement
patterns, and reproductive performance. However, the large size of
this tree squirrel allowed us to examine current ideas on the evolution
of sociality in squirrels in relation to body size. More than 2 dozen
undergraduate students have worked on this research project with me over
the years and several have produced senior theses from ecological investigations:
Habitat use and nest site location of Chiricahua fox squirrels--Nicole
Michel
Foraging behavior of male and female Chiricahua fox squirrels--Michele
Corse
Vigilance behavior of Chiricahua fox squirrels in relation to human activity--Mary
Kneeland
Interspecific competition,
seed removal, and population dynamics of mammalian granivores in
an oak-fir ecosystem
Dates:
1992-1995
Social organization
and habitat use of the collared peccary, Chiricahua National Monument,
AZ
Dates:
1994-1995
Conflict between
the sexes: social organization in tree squirrels, Univ. of Kansas
Dates:
1986-1992
Demography of fox
squirrels in a small urban woodlot, Southern Illinois University (w/W.D.
Klimstra)
Dates:
1983-1985
Teaching Experience:
Associate Professor of Wildlife
Science
Aug 2000-Present
Director of the Mt.
Graham Red Squirrel Monitoring Program
Wildlife
and Fisheries Science
School
of Renewable Natural Resources
University
of Arizona
Tucson,
AZ 8572
Courses:Wildlife
Management: Mammalian Species, Biogeography
and Conservation of the Sky Islands,
Applications
of Behavioral Ecology to Conservation Biology, Natural
Resources Ecology
Associate Professor of Biology
Aug 1997-Aug 2000
Assistant Professor of Biology
Aug 1992-Aug 1997
Department
of Biology
Willamette
University
Salem,
OR 97301
Courses:General
Ecology, Behavioral Ecology, Biological Diversity, Topics in Conservation
Biology,
Topics in Mammalogy, Field
Zoology, Vertebrate Zoology, Field Studies in Ecuador, Evolutionary
Biology, Life Forms 2: Biology of Higher Organisms, Senior Research Seminar,
Perspectives in Biology, Comparative Vert Morphology, Marine
Mammals, Principles of Biology, .
Courtesy Assistant Professor
of Biology
Mar 1995 - Present
Department
of Fish and Wildlife
Oregon
State University
Corvallis,
OR 97331
Courses:Ecological Consequences of Habitat Fragmentation
Lecturer
Department
of Biology
University
of Missouri-Kansas City Aug 1991 - May 1992
Kansas
City, MO 64110
Courses:Vertebrate Zoology, Evolution, Mammals of Kansas City, Urban Wildlife
Graduate Teaching Assistant
Department
of Systematics & Ecology
University
of Kansas
Jan 1986 - May 1991
Lawrence,
KS 66045
Courses:Laboratory
in Experimental Ecology, Honors Intro Biology Laboratory,
Honors Physiology Laboratory,
Animal Behavior Laboratory, Intro Biology Discussions
Adjunct Biology Faculty
Department
of Biology
Aug 1988 - Dec 1989
Kansas
City Kansas Community College
Kansas
City, KS 66012
Course: General Biology
Updated
12 February 2004
John
L. Koprowski